Bazzania trilobata

bz

bz
Shining Cliff Wood
Perhaps Derbyshire's most handsome liverwort, this species is very rare in the county and restricted to a few sites where it tends to be found in very small quantity.  Most populations are presumably relicts of pre-industrial times, when the species would have been more widespread and abundant.  At most of its current sites, B. trilobata occurs in deep crevices in block scree, where it was able to survive the widespread air pollution of the 19th and 20th centuries. 

The best population is probably in Shining Cliff Wood, where it is abundant around the edges of scree and is much more easily-seen than at other sites in the north of the county.

bz
 

 

 

Trichocolea tomentella

trc
below Stanage Edge
This beautiful liverwort is extremely rare in Derbyshire.  It has been recorded from a handful of sites, mostly in the High Peak, but has not been refound at any in recent years except for Whitwell Wood.

The tiny colony shown here was a new find in 2022.

Trichocolea is described in more detail in the Liverworts of Scotland section of the site.

 

 
 

Lepidozia reptans

lr
 

lr
 

lr
Hillbridge Wood NR

This common liverwort is a typical species of damp acidic woodland, especially on rotting logs.  It can easily be recognised by its bright green colour and pinnate branches with curved, lobed leaves. 

Like many liverworts, this species is much more abundant in the wetter west of Britain, and is relatively scarce in many areas of Derbyshire.

llr

 

 

Calypogeia muelleriana

cm
Hillbridge Wood NR

Although several Calypogeia species look similar, the acidic woodland habitat and rounded, entire leaf tips of this plant help identify the plant in these photos as C. muelleriana.  The underleaves of this species also have only a shallow notch and no extra lateral lobes - a useful feature to conclusively identify the plant. 

In all, six Calypogeia species are present in Derbyshire.

 

 

Calypogeia fissa

cpf
Grainfoot Clough, Derwent Valley
Named for the divided leaves, C. fissa is an easy plant to identify as it is the only Calypogeia (apart from C. arguta, below) with consistently bidentate leaf apices.  This character gives the leaves a notched appearance which is easily seen with a hand lens.

C. fissa grows in similar damp habitats to other Calypogeias and is a relatively common species.

 

 
 

Calypogeia arguta

ca
Bentley Brook, Lumsdale, Matlock
This common species of acidic habitats is described in the Bryophytes of Dorset section of the site.

 

 
 

Cephalozia bicuspidata

cb
 

cb
perianths, Padley Gorge (both photos)

This little liverwort can be found in most acidic habitats in Derbyshire, and is much more common than the other two widespread Cephalozia species (see below). 

The diagonally inserted 2-lobed leaves are a key diagnostic feature.  In winter, the very large perianths also become a conspicuous feature.

 

 
 

Cephalozia lunulifolia

cl

cl2
Padley Gorge
Cephalozias are small, mat-forming liverworts which can be tricky to identify in the field; the plant in these two photos was confirmed by microscopical analysis.

Key features of this species are its longitudinally-inserted leaves which often have a moon-shaped sinus (indentation) between the two lobes on each leaf.  Perianths are a good feature to look for as well - C. lunulifolia has untoothed female bracts. 

Only two other Cephalozia species are known from Derbyshire - the bog-dwelling C.connivens and the common C.bicuspidata (see above), which has diagonally inserted leaves.

 

 

Cephalozia curvifolia (syn Nowellia curvifolia)

nw
 

nwl2
Hillbridge Wood NR

nc2
Perianths, Mill Dale

Wet, rotting logs are the characteristic habitat of this widespread liverwort.  Although common in western Britain, this attractive species is not especially common in Derbyshire.

C. curvifolia can be recognised by its worm-like shoots and strongly concave leaves with two long pincer-like lobes.  The long, thin stems can completely cover rotting logs in woodland in favoured sites.  Another distinctive feature of the species is its crimson-red colour, although in deep shade red pigments can be completely absent.  The first photo shows a more typically coloured patch of plants. The photo below looks rather yellow as a result of the camera flash - the colour is actually a rich green with red tinges.

nw

nw
Via Gellia

 

 

Odontoschisma denudatum

odn
Hillbridge Wood NR
 
The most immediate distinguishing feature of this liverwort is the presence of abundant balls of yellow gemmae on the tips of the shoots.  Although some other liverworts have this feature, those species usually have toothed or nothced leaves (e.g. Barbilophozia attenuata - see below).  No British liverwort combines the features of yellow terminal balls of gemmae, rounded leaves and upright habit.

O. denudatum is primarily a western, upland species, and is a very rare plant in the Peak District.  It is only known from two sites in Derbyshire, and occurs in very small quantity at both.

 

 

Odontoschisma fluitans

odfl
White Path Moss, Burbage

This small liverwort is a bog specialist and is very rare in Derbyshire.  It closely resembles Gymnocolea inflata, which can grow in the same habitat and is much more common - microscopical confirmation is necessary, when the stem flagellae and conspicuous underleaves of Odontoschisma are easily seen.  The cells are also larger than in Gymnocolea.

Odontoschisma fluitans is only known from four 1km squares in the county - three on the Eastern Moors and one near Stanage Edge.

 

 

Barbilophozia attenuata

ba
 

ba
Hillbridge Wood NR

The abundant gemmae-bearing shoots of this fairly common liverwort make it perhaps the easiest British Barbilophozia to identify.

The scientific name comes from the tapered (attenuated) gemmae-bearing shoots.

 

 

Barbilophozia atlantica

ba
 

baWestend Valley, High Peak

Very similar to B.floerkii (below), this liverwort is fairly frequent on exposed acidic rock outcrops in the Peak District.

Although variable in colour, from yellowish, to green to brownish, at least some stems in a colony will usually be seen to have red patches of gemmae at the the tips - a feature always absent from B.floerkii.

 

 

Orthocaulis floerkii (syn. Barbilophozia floerkii)

bf
Ashop Clough, High Peak
brb
Wragg's Quarry, Eastern Moors

The two consistent features of this liverwort are the three-lobed leaves (not obvious from this photo) and the lack of gemmae.

Along with B. attenuata, this is the commonest Barbilophozia/Orthocaulis in Derbyshire, and in England as a whole.  It is found in acidic habitats, and is locally common in the Dark Peak on peaty rock ledges and rotting logs.

 

 

Barbilophozia barbata

bb
 

bh
Bar Brook, Eastern Moors

The rarest of Derbyshire's Barbilophozias, this species favours slightly base-enriched habitats, rather than the strongly acidic habitats occupied by its cousins.  It is seldom abundant, even in favoured sites, with sparse patches occurring on one or two boulders, while adjacent ones have none of the species present.

The easiest identification feature to see in the field is the way the young leaves at the tip of the shoots are pressed against each other, rather than spreading apart (as in other Barbilophozia species).  The leaves are clearly 3-lobed, making confusion with other small leafy liverworts unlikely.


bb

bb

 

 

Lophozia bicrenata

lb
 

lb
sporophytes, Tideswell Dale

A rare liverwort in Derbyshire, known from a few sites, L. bicrenata is a very small, close-leaved species, often noticeable by its orange-red gemmae which are borne abundantly at the shoot tips.  It grows on bare acidic soil where competition from other plants is reduced.

Microscopically, L. bicrenata is best identified by the thickened cell walls on the perianths.  These are the most reliable character to distinguish it from L. excisa, which grows in similar habitats and the two species sometimes grow together.  L. excisa has perianths with thin cell walls, purplish gemmae and smooth leaves.  Most sites for L. excisa in Derbyshire are from soil ledges on limestone outcrops, unlike L. bicrenata.

 

 

Lophozia excisa

le
Slitherstone Mine, SW of Castleton
More common in Derbyshire than L. bicrenata, L. excisa is described with that species, above.

 

 

Lophozia sudetica

lex
below Roych Tor, High Peak
ls
stem detail
Text coming soon.

ls
Gemmae

 

 

Oleolophozia perssonii

lp
Tearsall Quarry, near Winster
lg
Gemmae (mature are yellow with large oil bodies, immature are colourless with multiple small oil bodies)
This rare Lophozia species is a pioneer of strongly calcareous sites in Derbyshire, especially old limestone quarries.  It is known from scattered sites across the White Peak, but is always a special and unpredictable find.

In the field, this is an easy species to identify, as the stems always have dense clusters of dark brown gemmae at their tips.  Microscopically, the gemmae are fairly rounded and contain large, persistent oil bodies.  L. bicrenata could look fairly similar but is never found in calcareous sites and has thickened cell walls.  L. excisa grows in mildly-calcareous sites but has different perianth mouths, is paroicous and the leaves are inserted diagonally, rather than horizontally on the stems.  L. excisa also usually has only sparse gemmae (this is never the case in L. perssonii) which are purplish and contain small oil bodies.

 

 

Mesoptychia collaris (syn. Leiocolea alpestris)

la3
Monks Dale NNR
Despite its name, this small liverwort is by no means restricted to upland habitats.  It is a fairly frequent species of damp outcrops in limestone grassland in the White Peak, where it forms distinctive opaque pale green mats of scattered stems on bare soil.

It can be distinguished from the more common M. turbinata by its larger size and broad-based leaves, as well as the lack of underleaves.

 

 

Mesoptychia badensis (syn. Leiocolea badensis)

image coming soon!
This small liverwort of opn, stony soils is particularly common in Derbyshire as a colonist of old limestone quarries and mine sites, where it can be remarkably abundant, forming large, pure 'turfs' that stretch for metres.

M. badensis is usually an easy species to identify.  Like M. turbinata (below), it lacks underleaves and the two species are separated by how wide their leaf bases are.  Typical badensis has very widely-inserted leaves, although some stems can appear to have more narrow leaf bases which resemble M. turbinata.  The latter species has non-decurrent leaf bases.

 

 

Mesoptychia turbinata (syn. Leiocolea turbinata)

lt
Matlock Bath
Three Mesoptychia species are found in the Peak District, and this is perhaps the commonest of the three.

The best distinguishing feature of this lime-loving species is the narrow leaf base - unusual among British liverworts.  It is a very small species, more so than the other two, and is found on permanently wet limestone, such as seeping rock ledges and damp tufa.

lc2

 

 

Gymnocolea inflata

gy
 

gy
Westend Valley, High Peak

This liverwort of acidic habitats is especially common in upland gritstone areas where it forms dark patches of short, sprawling stems.  These have distinctive inflated perianths (hence the scientific name).

gy

 

 

Tritomaria quinquedentata

tqu2
 

tqu4
Coombs Dale SSSI

The most immediate feature of this fairly large liverwort is the way the leaves are split into three sharp lobes, with a long, curved back edge.  This asymmetric leaf shape is characteristic and distinguishes this species from otherwise similar Barbilophozia species.

Although mainly an upland species reaching its British south-eastern limit in the Peak District, it is not uncommon in the region and can be seen at many sites in the White Peak in calcareous grassland.

 

 

Jungermannia pumila

jp
Fernilee Reservoir
jp
Bentley Brook, Lumsdale, Matlock
As its name suggests, this is a small liverwort, and easily overlooked.  It favours damp ground.

Like many small Jungermannia species, fertile plants need to be located to help with identification.  J. pumila has a paroicous inflorescence (pairs of male bracts below a single pair of female ones) and lacks perigynia (sheaths surrounding the female inflorescence).  In the Peak District, at least, these features are conclusive.

This primarily northern and western species is fairly widespread in the Peak District, which represents the south-eastern limit of its main range in Britain.  A smaller, disjunct range exists in Kent and Sussex.

 

 

Jungermannia eucordifolia (syn. Jungermannia exsertifolia ssp. cordifolia)

ja
Mill Dale, Ladybower
One of the rarer Jungermannias in Derbyshire, this northern species is restricted to a few moorland streams in the High Peak.

The species is described in more detail in the Liverworts of Ben Lawers NNR section of the site.

 

 

Solenostoma sphaerocarpa

j
 

js
Westend Valley, Dark Peak

Although rather similar to S. gracillimum (below), this very small liverwort can be often be recognised by its liking for wet habitats, such as dripping rock ledges or emergent stones in and by upland streams.

Unlike its close cousin (which favours drier, disturbed habitats) it is a paroicous species, with male and female parts on the same stem.  It also lacks the red-coloured perianth and strongly-bordered leaves of S. gracillimum, although a hand lens is essential for viewing these features.

Both species are more or less equally common in northern Derbyshire.

 

 

Solenostoma gracillimum

jg
Brampton Moor

jgr
Perianths, Ramsley Reservoir, Eastern Moors

In Derbyshire this diminutive liverwort of disturbed, bare, acidic soils is restricted to the moorland areas of the Peak District.

The classic diagnostic feature for this species is the margin of enlarged cells along the leaf borders, but these can be hard to see, given the tiny (less than 1mm) width of the leaves - a x20 hand lens is essential.  Habitat also gives a good hint for identifying this species in the county.

The second photo shows the striking red perianths, which can be very abundant in some colonies.

 

 

Solenostoma hyalinum (syn. Jungermannia hyalina)

hyal
Gregory Mine, Ashover
This small species is very rare in Derbyshire, and currently known from a single site, where it is locally abundant on a damp soil bank.

S. hyalinum is one of three Derbyshire species that routinely have purple rhizoids on the stems - this is a useful character.  Confirmation of its identify, though, requires examining perianths on fertile material.  Sterile plants can be suspected but not confirmed.  S. hyalinum is a dioicous species and the female stems bear perianths with long cells at the mouth and well-developed perigynia.  Sometimes S. paroicum can be dioicous and strongly resemble S. hyalinum but it has larger cells.

 

 
 

Jungermannia atrovirens

ja
 

jaMill Dale

This species is the only calcicolous Jungermannia in Derbyshire, where it forms small dark green stems on wet rock faces or damp grassy slopes.

A key morphological feature is the dioicous inflorescence, with male and female plants being separate.  Other dioicous Jungermannias/Solenostomas in Derbyshire are J. atrovirens, S.gracillimum (see above) and S.hyalinum, but these avoid calcareous habitats, and so are unlikely to cause confusion in the limestone dales where J. atrovirens is common.

 

 

Nardia scalaris

nr
 

nr
Westend Valley, High Peak

This liverwort is described in the Liverworts of Ben Lawers NNR section of the site.

nrd
Wragg's Quarry, Eastern Moors

 

 

Nardia compressa

nc
Big Moor, Eastern Moors
N. compressa is an attractive liverwort typical of acidic, rocky moorland streams in the High Peak in Derbyshire.  Here it can be a dominant species.  Identification is straightforward - the stems have leaves pressed against them giving a thin appearance, quite unlike most British liverworts.  This is the origin of the scientific name.

Like many liverworts, the Peak District represents the south-eastern limit of this species' British distribution, apart from a small, disjunct population in East Sussex.

nc
 

 

 

Diplophyllum albicans

da
perianths, Ladybower Wood, Derwent Valley
Another common liverwort in Derbyshire, this species is described in the Liverworts of Ben Lawers NNR section of the site.

 

 

Scapania compacta

sc
 

sc

sc
Tideswell Dale

One of the rarer Scapanias in Derbyshire, this attractive and distinctive species is only abundant in Tideswell Dale, where it grows on an exposed basaltic outcrop.

The closely-pressed, equal-sized front and back leaf lobes give this liverwort a neat, symmetrical appearance.

sc

 

 

Scapania undulata

su
Fertile plants with perianths, Ashop Clough
This common liverwort is widespread in Derbyshire.

I've described the species in the Liverworts of other areas of Scotland section of the site, but have included this photo as it shows the abundant perianths of a fertile specimen.

 

 

Scapania aspera

sc

sca
Coombs Dale SSSI
The most lime-loving lowland British Scapania, this species is common in the White Peak, where it sometimes grows luxuriantly in shaded woodland among limestone rubble and on crags.  In the limestone dales this is often the only Scapania present.

When viewed closely, the front leaf lobe extends well across and runs slightly down the stem.  Where present, branches arise from the back of stem - a feature almost unique to this member of the genus.  The most similar species, S. aequiloba, is not found in Derbyshire. 


sc

 

 

Scapania nemorea

sn
 

scn
Hillbridge Wood NR

sn
Bar Brook, Eastern Moors

This common woodland Scapania is described in the Bryophytes of Cumbria section of the site.

Along with S. undulata (above), this is by far the commonest Scapania in acidic upland areas of Derbyshire.


sn

 

 

Scapania umbrosa

sc
Padley Gorge
Scapania umbrosa is a great rarity in Derbyshire, despite its rather mundane habitat requirements.  Its typical habitat here is as a pioneer species on sandy gritstone boulders in woodland.

In the field, the most apparent feature of this small species is the abundant red gemmae.  Microscopically, the leaves are seen to have very pointed lobes with sharply-toothed margins and the gemmae have very thin cell walls.

 

 

Scapania gracilis

sc
 

sc

sca
Ladybower Wood, Derwent Valley

In northern and western Britain, this species is common in damp woodland.  In Derbyshire, it is very rare and more or less on the edge of its national distribution.

S. gracilis often has green gemmae present, and this, combined with its acidic woodland habitat, gives a good hint as to its identity.  Microscopically, the leaves usually have teeth at their bases - an unusual feature among Scapanias.  In woodland, S. nemorea and S. undulata can sometimes have this too, but they differ in leaf shape and/or gemmae colour and most of their leaves will lack the basal teeth.

scg

 

Scapania irrigua

sci
 

sc
Wragg's Quarry, Eastern Moors

Another rare Scapania in Derbyshire, this species is characteristic of open, gritty habitats like old quarry floors.  Derbyshire's largest population is probably at Wragg's Quarry near Matlock, where it is a dominant species covering large areas in and around a marshy depression in the quarry floor.

Under a microscope, the leaves can be seen to have pointed apices with few or no teeth and a fairly weakly-curved keel.  The smaller (antical) lobe is squarrose, giving the shoots a more three-dimensional appearance than some Scapania species.  In my experience, gemmae are not common here.

sc

 

 

Lophocolea bidentata

lc
 

lc
Hillbridge Wood NR

The commonest Lophocolea in the limestone dales of the Peak District, this woodland species can be recognised by its regularly and strongly toothed (two-pronged) leaves.

It is an abundant liverwort of shaded, damp drystone wall communites, old limestone quarries and among rocks and tree roots on the woodland floor.  It can also be found on rotting logs in woodland (as in the second photo).

 

 

Pedinophyllum interruptum

pi
 

pi
Chee Dale SSSI

This plant is one of Derbyshire's rarest liverworts.  Its only English occurrence south of the Yorkshire Dales is in the Chee Dale area, where it grows on wet, wooded, limestone crags, although historically it was recorded from Matlock Bath as well.

It can be recognised by its prostrate shoots which taper towards the tip, its rounded leaves and the sparse appearance of the shoots due to the midline of the stems being free of leaves.

 

 

Plagiochila asplenioides

pa
Chee Dale SSSI

pa
Rose End Meadows, Cromford

Britain's second commonest Plagiochila is fairly easy to recognise, and often catches the eye as a distinctly large liverwort.

It has rounded leaves like P. porelloides (below), but is nearly always larger, and is a pale green or yellowish-green colour, rather the dark green of its cousin.

pa

 

 

Plagiochila porelloides

pl
Coombs Dale SSSI
Resembling a miniature version of P. asplenioides, this is the commonest Plagiochila in limestone areas of the Peak District (although it also comonly grows in acidic areas), and the commonest member of the genus nationally.

It can be recognised by its broadly rounded, unlobed leaves with toothed margins.  Fertile plants are rare in the Peak District.

 

 

Chiloscyphus polyanthos

chl
Bentley Brook, Matlock

A common and attractive liverwort of frequently-submerged, acidic rocks in streams, this species is easy to encounter in Derbyshire.  It is also common in moorland flushes and springheads.

The broad-based, rounded leaves and more-or-less aquatic habitat will identify this species in most cases.

 
 

 

 

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